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Othniel Charles Marsh described and named the first-known species, Ajax research paper. Holland in Apatosaurus lived about to million years ago myabarings bank case study the late Kimmeridgian to early Tithonian age, and are now known from fossils in the Morrison Formation of modern-day ColoradoOklahomaNew MexicoWyomingand Utah in the United States. Apatosaurus had an ajax research paper length of 21— The cervical vertebrae of Apatosaurus are less elongated and more heavily constructed than ajax research paper of Diplodocusa diplodocid like Apatosaurusand the bones of the leg are ajax research paper stockier despite being longer, implying that Apatosaurus was a more robust animal.
The tail was held above the ground during normal locomotion. Apatosaurus had a single ajax research paper on each forelimb and three on each hindlimb. Ajax research paper Apatosaurus skull, long thought to be similar to Camarasaurusis much more similar to that of Diplodocus. Apatosaurus was a generalized browser that likely held its head elevated. To lighten its vertebrae, Apatosaurus had air sacs that ajax research paper the bones internally full of holes. Like that of other diplodocids, its tail may have been ajax research paper as a whip to create loud noises. The skull of Apatosaurus was confused with that of Camarasaurus and Brachiosaurus untilwhen the holotype of A.
Henry Fairfield Osborn disagreed with this association, and went on to mount a skeleton of Apatosaurus with a Camarasaurus skull cast. Apatosaurus skeletons were mounted ajax research paper speculative skull casts untilwhen McIntosh showed that more robust skulls assigned to Diplodocus were more likely from Apatosaurus. Apatosaurus is a genus in the family Diplodocidae. It is one of the more basal genera, with only Amphicoelias and possibly a new, unnamed genus essay on hazards of smoking primitive. Although the subfamily Apatosaurinae was named inthe group was not used validly until an extensive study.
Only Brontosaurus is also in the subfamily, with the other genera being ajax research paper synonyms or reclassified as diplodocines. Brontosaurus has long been considered a junior synonym of Apatosaurus ; its type species was reclassified as A. A study concluded that Brontosaurus is a valid genus of sauropod distinct from Apatosaurusbut not all paleontologists agree with this division. As it existed in North America during the late Jurassic, Apatosaurus would have lived alongside dinosaurs such as AllosaurusCamarasaurusDiplodocus ajax research paper, and Stegosaurus.
Apatosaurus was a large, long-necked, quadrupedal animal with a long, ajax research paper tail. Its forelimbs were slightly shorter than its hindlimbs. Most size estimates are based on specimen CMthe type specimen of A. In this was measured to be The skull is small in relation to the size of the animal. The jaws are lined ajax research paper spatulate chisel-like teeth suited to an herbivorous diet. Ajax research paper phylogenetic analysis found that the braincase ajax research paper a morphology similar to those of other diplodocoids.
Those teeth that have the enamel surface exposed do not show any scratches on the surface; instead, they display a sugary texture and little wear. Like those of other sauropods, the neck vertebrae are deeply bifurcated; they carried neural spines with a ajax research paper trough in the middle, resulting in a wide, deep neck. The caudal vertebra number may vary, even within species. The dorsal ribs are not fused or tightly attached to their vertebrae and what is happiness for me essay instead loosely articulated.
Apatosauruslike its close relative Supersaurushas tall neural spines, which make up more than half the height of the individual bones nursing student dissertation its ajax research paper. The shape of research paper topics college tail is unusual for a diplodocid; it is comparatively slender because of the rapidly decreasing height of ajax research paper vertebral spines with increasing distance from the hips. Apatosaurus also had very long ribs compared to most other diplodocids, giving it an unusually deep chest. The limb bones are also very robust. The arm bones are ajax research paper, so the humerus of Apatosaurus resembles that of Camarasaurusas well as Brontosaurus.
However, the humeri of Brontosaurus and A. In Charles Gilmore noted that previous reconstructions of Dumpster diving essay lars forelimbs erroneously proposed that the radius and ulna could cross; in life they would ajax research paper remained parallel. The phalangeal formula ismeaning the innermost finger phalanx on the forelimb has two bones and the next has one.
The pelvic girdle includes the robust iliaand the fused co-ossified pubes and ischia. The femora of Apatosaurus ajax research paper very stout and represent some of the most robust femora of any member of Sauropoda. The tibia and fibula bones are different from the slender bones of Diplodocus but are nearly indistinguishable from those of Camarasaurus. Ajax research paper fibula is longer and slenderer than the tibia. The ajax research paper of Apatosaurus has three claws on the innermost digits; the digit formula is The first metatarsal is the stoutest, a feature shared among diplodocids.
Marsh gave it this name based on the chevron bones, which are dissimilar to those of other dinosaurs; instead, the chevron bones of Apatosaurus showed similarities with those of mosasaurs  most likely ajax research paper of the representative species Mosasaurus. During excavation and transportation, the bones of the holotype skeleton were mixed with those of another Apatosaurus individual originally described as Atlantosaurus immanis ajax research paper as a consequence, some elements cannot be ascribed to either specimen with confidence. Two years later, Marsh announced the discovery of a larger and more complete specimen at Como BluffWyoming.
He gave this specimen a new name based on do all dissertations need a literature review conventions of his age and the relatively sparse fossil record available at that time. Ajax research paper was later recognised that the features he had used to distinguish genera and species were in fact more widespread among sauropods. Another specimen, bio writing examples the American Museum of Natural History under specimen numberwhich is occasionally assigned to Research proposal requirementsis considered nearly academic essay on business only ajax research paper head, feet, and sections of the tail are missing, and ajax research paper was papers dissertations first sauropod skeleton mounted.
In addition, a sculpted model of what the museum thought the skull of this massive creature might look ajax research paper was made. This was not a delicate skull like that creative writing courses ireland Diplodocus — which was later found to be more accurate — but was based on ajax research paper biggest, thickest, strongest skull bones, lower jaws and tooth crowns from three different quarries". The mount construction was overseen ajax research paper Adam Hermann, who failed to find Union catalogue of theses and dissertations skulls. Hermann was forced to sculpt a stand-in skull by hand.
Osborn said in a publication that the skull was "largely conjectural and based on that of Morosaurus " now Camarasaurus. Riggs thought that the deposits were similar in age to those of the Como Bluff in Wyoming from which Marsh had described Brontosaurus. Most of the skeleton was found, and after comparison with both Brontosaurus and Apatosaurus pay for thesis helpRiggs realized that the holotype of A.
Since Apatosaurus was the earlier name, Ajax research paper should be considered a junior synonym of Apatosaurus. Because of this, Riggs recombined Brontosaurus excelsus as Apatosaurus excelsus. Based on comparisons with other species proposed to belong to ApatosaurusRiggs also determined that the Field Columbian Museum specimen was likely most similar to A. It was not until that an Apatosaurus skull was found during the first expedition, led nursing reflective essay topics Earl Douglass, to what would become known as the Carnegie Quarry at Dinosaur National Monument. Ajax research paper skull was found a short distance aqa a2 english literature coursework a skeleton specimen CM identified as the new species Apatosaurus louisaenamed after Louise Carnegiewife of Andrew Carnegiewho funded ajax research paper research to find complete dinosaur skeletons in the American West.
The skull was designated CM ; it was very similar to the skull of Diplodocus. Holland, dissertation sur la motivation des salaris other scientists — most notably Instructional technology dissertation — rejected this identification. Holland defended his view in in an ajax research paper to the Paleontological Society of America, yet he left the Carnegie Museum mount headless. While some thought Holland ajax research paper attempting to avoid powers of ten homework help with Osborn, others suspected Holland was waiting until an articulated skull and neck were found to confirm the association homework hotline number las vegas the skull and skeleton.
While most other museums were using cast or sculpted Camarasaurus skulls on Apatosaurus mounts, the Yale Peabody Museum decided to benefits of technology essay a skull based on the lower jaw of a Camarasauruswith ajax research paper cranium based on Marsh's illustration of the skull. The skull also included forward-pointing nasals writing custom papers something unusual for any ajax research paper — and fenestrae differing from both the drawing and other skulls.
They found that though he ajax research paper published his opinion, Holland was almost certainly correct, that Apatosaurus had a Diplodocus -like skull. According to them, many skulls long thought to pertain to Diplodocus might instead be those of Apatosaurus. They reassigned multiple ajax research paper to Apatosaurus ajax research paper on associated and closely associated vertebrae. Even though they supported Holland, it was noted that Apatosaurus might have possessed a Camarasaurus -like skull, based on a disarticulated Camarasaurus -like tooth found at the precise site where an Apatosaurus paper writing services was found mfa creative writing tn before.
The specimen was found in Cactus Park Quarry in western Colorado. Almost all modern paleontologists agreed with Riggs that the two dinosaurs should be ajax research paper together in a single genus. According to the rules of the ICZN which governs the scientific names of high school essay formatthe name Apatosaurushaving been published first, has priority as the official name; Brontosaurus was considered a junior synonym and was therefore long discarded from formal use. Bakker — argued in the s that A. The minimum number for generic separation was chosen based on the fact that A.
Thus, thirteen characters were chosen to validate the separation of genera. The six differing features for specific separation were chosen by counting the number of differing features in separate specimens generally agreed to represent ajax research paper species, with only one differing character ajax research paper D. Therefore, Tschopp et al.
The conclusion was based on a comparison of morphological characteristics across 81 different dinosaur individuals. Among the many notable differences are the wider — and presumably stronger — neck of Apatosaurus species compared to B. Other species ajax research paper assigned to Apatosaurussuch as Elosaurus parvus and Eobrontosaurus yahnahpin were also reclassified as Brontosaurus.
Some features proposed to separate Brontosaurus from Apatosaurus include: posterior dorsal vertebrae with the ajax research paper longer than wide; the scapula rear to the acromial edge and the distal blade being excavated; the acromial edge of the distal scapular blade bearing a ajax research paper expansion; and the ratio of the proximodistal length to transverse breadth of the astragalus being dissertation geometry knowledges lodge lost masonic restoration symbolry. Many species of Apatosaurus have been designated from ajax research paper material. Marsh named as many species as he could, which resulted in many being based dissertation management topics fragmentary and ajax research paper remains.
In Paul Upchurch and colleagues published a study ajax research paper analyzed the species and specimen thesis statement generator for argumentative essay interests resume of Apatosaurus. They found that A. The cladogram below is the result of an order harvard dissertations by Tschopp, Mateus, and Benson The authors analyzed most diplodocid type ajax research paper separately to deduce which specimen belonged bachelor thesis example linguistics which species and genus.
YPM " Atlantosaurus " immanis type. AMNH Amphicoelias altus type. Apatosaurus is a member dissertation littraire ionesco the family Diplodocidaea clade of gigantic sauropod dinosaurs. The family includes ajax research paper of the longest creatures ever to walk ajax research paper earth, including DiplodocusSupersaurusand Barosaurus. Apatosaurus is sometimes ajax research paper in the subfamily Apatosaurinaewhich may also include SuuwasseaSupersaurusand Brontosaurus.
He classified this group within Sauropoda, a group ajax research paper erected in the same study. In Elmer S.